CHANDLER'S CROSS, England – The juggling and adjusting continues.

And at least to some extent, the Jaguars could be dealing with more injuries at wide receiver, too.

Marqise Lee and Cecil Shorts III, a pair of Jaguars wide receivers who have missed extensive time with injuries this season, appeared on the team's injury report Wednesday after experiencing issues during the team's first full workout of the week.

Lee, a rookie second-round selection in the 2014 NFL Draft, sustained an ankle injury in the team's workout at the Grove Hotel practice field. Lee was listed as limited on the injury report.

Cecil Shorts III, a veteran starter, experienced soreness in his hamstring during practice Wednesday, and also was listed on the Wednesday injury report as limited. Shorts missed two preseason games and three regular-season games with hamstring issues this season.

"We'll see how it goes – I know with Cecil … he knows his body so well," Head Coach Gus Bradley said on Jaguars.com Live Wednesday afternoon as the Jaguars (1-8) prepared to play the Dallas Cowboys (6-3) at Wembley Stadium in London Sunday at 1 p.m.

"It got to that point where it was time. He'll get treatment and we'll see how he is (Thursday)."

Lee, a starter in the first two games of the regular season and against the Dolphins in Week 8, has caught 12 passes for 128 yards this season, missing three games with a hamstring injury in September and early October. He said earlier in the day he hadn't lost confidence in the first nine games, and Bradley had said while meeting with the media this could be a big week for Lee for the team to see how he practiced.

Bradley said on jaguars.com that Lee's spirits were good after the injury.

"It scared him more than anything," Bradley said.

Shorts, a starter in five games this season – including the last three – has 28 receptions for 256 yards and a touchdown in six games played.

The news regarding Shorts and Lee came the same day the Jaguars were dealing with injuries at two other positions, with familiar faces possibly returning to the lineup as a result.

Brandon Linder, a rookie offensive guard who has earned praise with his play in the first nine games, did not practice with a shoulder strain sustained against the Cincinnati Bengals last week. Bradley said he doesn't expect Linder to practice Thursday, either.

Jacques McClendon, a starter at center in the first two games of the regular season, could start at right guard Sunday if Linder can't play, Bradley said. McClendon guard during the preseason and has worked as the backup there during the season.

Bradley said rookie Tyler Shatley also is a possibility at right guard. The Jaguars liked Shatley coming out of the preseason enough that they kept him on the roster for fear of losing him to another team if they tried to keep him on the practice squad.

Defensively, the Jaguars are once again dealing with injuries at linebacker.

"It's a little bit concerning," Bradley said.

Rookie Jeremiah George, who played impressively early against Cincinnati Sunday in his first NFL start, missed practice Wednesday with a high-ankle sprain sustained in that game. He also is not expected to practice Thursday, Bradley said.

J.T. Thomas, who started at middle linebacker against Miami two weeks ago, likely will return to the middle with George out. Paul Poslusnzy started the first seven games of the season in the middle.

"J.T. gives us great flexibility," Bradley said. "He's played Mike. He's played Otto. Now it's back to Mike. We feel good about that."

Said Thomas, "Anyway I can add value to the team, I'm ready for it."

Thomas added that he will continue to call defensive signals and will continue to wear the sideline-to-player communication device. He wore that in the Otto role last week when George started inside.

"I was calling all the plays last week and I'll be calling them this week as well," Thomas said. "Everybody in our room is going to have to be better. This is going to be group effort. This defense is going to take all 11 of us. If I'm the guy who can put everybody on the same page, so be it."

With Thomas inside, Dekoda Watson likely will start at the Otto position.

Watson missed training camp and the preseason with a groin injury, and while he was signed as an unrestricted free agent to play the newly-created Otto position he has entered just one game as the No. 1 Otto. That was two weeks ago against Miami.

He missed last week's game with a hamstring injury while Thomas played the Otto. Thomas also started Weeks 4-7 at the Otto, with second-year veteran LaRoy Reynolds starting Weeks 1-3 there. Watson worked full on Wednesday.

"Dekoda seems like he's coming back," Bradley said. "I know he was nursing some hamstring injuries last week. We felt at that time that a week off would enable him to come back sooner.  Hopefully, he has.  I know he feels much better.

"With him being at Otto, we've got guys who have played a couple of games now."

Also around the Jaguars Wednesday:

*Cornerback Aaron Colvin (knee) worked full for the first time with the Jaguars. A rookie from Oklahoma, he has been on the Physically Unable to Perform list with a torn anterior cruciate ligament sustained at the Senior Bowl. The Jaguars have until next week to activate him or place him on season-ending injured reserve. …

*Quarterback Blake Bortles was listed on the injury report with a left wrist sprain. He said earlier in the day he's 100 percent. He worked full Friday. …

*Tight end Marcedes Lewis (ankle) practiced full, but can't play Sunday because he remains on the injured reserve-designated to return list. He is eligible to return November 23 at Indianapolis. …

*Safeties Josh Evans and Sherrod Martin practiced limited with shoulder injuries sustained against Cincinnati. …

*Defensive end Andre Branch (groin) did not practice and did not make the trip to London. He is expected to be out at least another month. …

*Bradley said newly-signed Khairi Fortt could be active Sunday as a backup linebacker/special teams player. "I think he has the traits that we're looking for, especially at the Otto position," Bradley said of Fortt, who signed off the Cincinnati practice squad Monday. …

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