JACKSONVILLE – Paul Posluszny's status for Sunday remains in question. Really, really in question.

Dan Skuta's status is in question, too, but the Otto linebacker's health has improved in recent days and he has a chance to play Sunday when the Jaguars play the Indianapolis Colts.

Skuta's return would be a positive for a defense that could be without Posluszny, the team's starting middle linebacker who missed the Jaguars' first practice of the week Wednesday after undergoing hand surgery Tuesday.

Posluszny sustained a fractured hand Sunday against Tennessee this past Sunday, but Jaguars Head Coach Gus Bradley on Wednesday did not rule out Posluszny playing against the Colts.

"We're waiting to see about his availability," Bradley said of Posluszny as the Jaguars (4-8) prepared to play the Colts (6-6) at EverBank Field Sunday at 1 p.m.

"We do know if he played he would play with a cast on."

Posluszny, a defensive captain, has started 11 of 12 games this season. He missed a 38-31 Week 5 loss to the Tampa Buccaneers in which the Jaguars allowed 183 yards, their second-most rushing yards allowed this season.

"The mike (middle) linebacker makes a lot of the (defensive) calls, and does a lot of checks, so that part of it (is important)," Bradley said. "It'd be great to have him out there. I think the big thing would be just the challenge of seeing what he could do with a club (on his hand), if that's what it is."

Bradley said Posluszny could be a game-time decision Sunday, adding that Posluszny was traveling to Jacksonville Wednesday and he had not discussed the situation with the nine-year veteran.

Skuta – like Posluszny a key to the Jaguars' run defense – worked limited in practice Wednesday with a groin injury. He has played the Otto position in nine of 12 games this season, missing three of the last four.

Bradley said Skuta's playing status also could be uncertain until Sunday.

"He did pretty good," Bradley said of Skuta in practice Wednesday.

The Jaguars received positive news on two injured players Wednesday, with wide receiver Rashad Greene and cornerback Aaron Colvin working full. Greene left Sunday's game with a shoulder injury, and Colvin played through a heavily-taped sore ankle. Neither was listed on the injury report.

Wide receiver Allen Hurns, who missed Sunday's game with a concussion sustained the previous week against San Diego, also worked full Wednesday. He never had missed an NFL game in two seasons before missing Sunday.

Safety Sergio Brown (hamstring), tight end Nic Jacobs (hamstring) and Skuta were listed as limited Wednesday. Brown and Jacobs both played against Tennessee Sunday.

Safety Johnathan Cyprien (stinger) did not practice Wednesday, with defensive end Chris Clemons (veteran day off), defensive end Ryan Davis (knee), defensive tackle Roy Miller (knee), Posluszny and wide receiver Bryan Walters (back) also not practicing. All of those players played Sunday.

Bradley said he expects Cyprien to play Sunday, and that Davis could be day-to-day.

Bradley on Wednesday discussed the overall play this season of Miller, a nose tackle who signed with the team as an unrestricted free agent in the 2013 offseason and who has played at a high level this season. Miller's performance is a big reason the Jaguars are ranked 13th in the NFL against the run.

Miller played through a shoulder injury in 2013 before missing the final two games, then missed the final two games last season with a knee injury.

"First and foremost, he's healthy," Bradley said. "He's always been a good player, but he's playing healthy this season. He has had a very good year. You talk about consistency; it's game in and game out."

Bradley said the Jaguars had a "good work day" Wednesday, with the focus for the week being on improving tackling and other fundamentals. The Jaguars missed 16 tackles in a 42-39 loss to Tennessee Sunday in which the Titans produced five offensive touchdowns and 467 total yards.

"It was a very competitive day again," Bradley said, adding, "We talk about it: it's about us. But this week it really is. We have to get some things corrected and that has been the emphasis for us: tackle better, use our fundamental principles, cover better – and then let's see."

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